Collybia effusa


ヒロハアマタケCollybia effusa Har. Takah., Mycoscience 42: 339-345, 2001        
 Figs. 1, 2 
  肉眼的特徴: 傘は径 30-45 mm, 最初半球形で縁部は内側に巻き, のち饅頭形からほぼ平
開し, しばしば中央部がやや凹み, 最初平坦であるがまもなく放射状に走る明瞭な扇状の溝線
を表し, 最初白色粉状, のち次第に平滑になり, 幼時赤褐色のち縁部に向かって淡色になる. 
肉は相対的に薄く (傘の中央部において厚さ 3mm 以下), 白色, 特別な味や臭いはない. 柄
は 30-55×1-3 mm, ほぼ上下同大, 中心生, 痩せ型, 中空, 白色, 全体に著しい粉状を呈し, 基
部は白色綿毛状菌糸体に被われる. ヒダは上生〜やや垂生し, 疎, 柄に到達するヒダは 13-
18, 幅 5 mm以下, 白色または時に緑黄色〜淡緑色を帯び, 縁部は平坦, 縁取りを欠く. 胞子
紋は白色.
  顕微鏡的特徴: 担子胞子は 7-8.5×3.5-4μm [Q = length/breadth: 2.1], 楕円形, 平坦, 無
色, 非アミロイド, 薄壁. 担子器は 28-38×7-8μm, こん棒形, 4胞子性. 偽担子器はこん棒形.
縁シスチジアは 22-60×4-8μm, 群生し, 類円柱形, 平坦または複数の瘤状突起を有し, 無
色, 薄壁. 側シスチジアはない. 子実層托実質は不規則に配列する; 菌糸は傘実質と共通す
る. 傘の表皮は平行菌糸被で, 匍匐性の菌糸が不規則に配列し, 瘤状突起に著しく被われるこ
とはない; 菌糸は幅 3-8μm, 錯綜し, 円柱形, 平坦もしくは少数の瘤状〜短指状突起に被わ
れ, 褐色の粒状色素に被覆され, 薄壁, 時に隔壁にクランプを持つ. 柄シスチジアは 35-60×6-
10 μm, 群生し, 屈曲した円柱形, 平坦もしくは少数の瘤状分岐物を有し, 無色, 薄壁. 実質の
菌糸はしばしば隔壁にクランプを有する.
  供試標本: KPM-NC0005067 (基準標本), Mt.Takao, Hachiouji-shi, Tokyo, 15 Jul. 1999, coll.
W. Ikeda & H.Takahashi; the same place, 7 Jun. 1999, coll. W.Ikeda & H.Takahashi; the same
place, 5 Nov. 1999, coll. W.Ikeda & H.Takahashi; the same place, 29 Jun. 2000, coll. H.
Takahashi.
  コメント: 子実体はモリノカレバタケ属としては大型で, しばしば群生し, 明瞭な放射状の溝線
を表す褐色の傘と, 白色または緑黄色の粗いヒダ,そして白色, 粉状の柄を有する. 東京都高
尾山のスギ及びムラサキシキブの落枝上に発生. 標本は神奈川県立生命の星・地球博物館
の標本庫(KPM)に登録, 収蔵されている.
  本種は, 傘表皮の菌糸の特徴, 胞子の生化学的性質, 及び生態に基づき, Antonin and
Noordeloos (1997) の定義による Gymnopus 属に置くべき種類と考えられる.

Collybia effusa Har. Takah., Mycoscience 42: 339-345, 2001         Figs. 1, 2 
  Pileus 30-45 mm in diam, at first hemispherical with involute margin, then plano-convex
to plane, often with slightly depressed center, at first smooth but soon radially plicate to
sulcate-striate almost to the disk, at first white pruinose, glabrescent in age, evenly colored
reddish brown (8D7-8 to 9D7-8) when young, then paler toward the margin. Flesh thin (up
to 3 mm), white; odor and taste not distinctive. Stipe 30-55×1-3 mm, almost equal, central,
slender, terete, hollow, white, pruinose to flocculose overall; base covered with white
tomentum. Lamellae adnate to subdecurrent, distant (13-18 reach the stipe), up to 5 mm
broad, moderately thin, white or sometimes yellowish green (30A5) to light green (30A6);
edges even, concolorous.
  Spore print pure white. Basidiospores 7-8.5×3.5-4 μm [Q = length/breadth: 2.1],
ellipsoid, smooth, colorless, inamyloid, thin-walled. Basidia 28-38×7-8 μm, clavate, four-
spored. Basidioles clavate. Cheilocystidia 22-60×4-8 μm, abundant, subcylindric, smooth
or sometimes with a variable number of outgrowths, colorless, thin-walled. Pleurocystidia
absent. Hymenophoral trama irregular; element hyphae similar to those of the pileitrama.
Pileipellis a layer of repent, branched hyphae, not distinctly diverticulate, not radially
arranged; constituent hyphae 3-8 μm wide, interwoven, cylindric, smooth or with a few
warty or finger-like protuberances, encrusted with granules of brown pigment, thin-walled,
occasionally with clamped septa. Hyphae of pileitrama 3-8 μm wide, interwoven, cylindric,
with smooth, slightly thickened walls up to 0.5 μm, colorless, inamyloid, occasionally with
clamped septa. Stipitipellis a cutis of parallel, repent hyphae 3-10 μm wide, cylindric,
smooth, thin-walled, colorless, inamyloid, occasionally with clamped septa; caulocystidia 35-
60×6-10 μm, abundant, often flexuous, cylindric, smooth or with a few knob-like
diverticula. Stipe trama composed of longitudinally running, cylindric hyphae 3-8 μm wide,
unbranched, smooth, slightly thick-walled (up to 0.5 μm thick), colorless, inamyloid,
occasionally with clamped septa.
  Known distribution: Japan (Tokyo).
  Habitat: Solitary to caespitose, on dead twigs of Cryptomeria japonica and Callicarpa
japonica, from June to November, not common.
  Specimens examined: KPM-NC0005067 (holotype), Mt.Takao, Hachiouji-shi, Tokyo, 15 Jul.
1999, coll. W. Ikeda & H.Takahashi; the same place, 7 Jun. 1999, coll. W.Ikeda & H.Takahashi;
the same place, 5 Nov. 1999, coll. W.Ikeda & H.Takahashi; the same place, 29 Jun. 2000, coll.
H.Takahashi.
  Japanese name: Hiroha-amatake (first collected and named by Mr.Minoru Aoki).
  Notes: This species is characterized by its distinctly sulcate-striate pileus, the
conspicuously white pruinose to flocculose stipe, distant lamellae, and its habitat on dead
twigs. The non radial arrangement of its pileipellis elements without a distinct Rameales-
structure (Dryophila-structure) suggest that this species belongs in the section Levipedes 
(Fr.) Quel. (Antonin and Noordeloos, 1997; Halling, 1983; Singer, 1986), where it appears to
be closely related to Collybia subcyathiformis (Murrill) Pegler from East Africa and
neotropical regions (Pegler, 1977). The latter species differs in having a dark brown pileus, a
conspicuously elongated stipe, and habitat on dead leaves. Collybia effusa has superficial
resemblance to two taxa of the section Vestipedes (Fr.) Quel. because of a strongly radially
sulcate-striate pileus, viz. Collybia multijuga (Berk. & Broome) Sacc. from Sri Lanka (Pegler,
1986), and Collybia allegretii (De Seynes) Pegler from East Africa (Pegler, 1977). These
species differ from C.effusa in forming a cutis of radially arranged hyphae in the pileipellis
(Confluens-structure). North American Collybia dysodes Halling (Halling, 1983), which
belongs to the section Sunfumosae Singer ex Halling, has a conspicuously plicate-sulcate
pileus and distant lamellae. Collybia dysodes, however, differs in having fetid odor, a reddish
brown stipe, adnate to adnexed, light brown lamellae, and a cutis of distinctly diverticulate
elements in the pileipellis.


Fig. 1.Collybia effusa.
A. Basidiospores. B. Basidium and basidiole. C. Cheilocystidia. D. Elements of the
pileipellis. E. Caulocystidia. Scales: 10µm. All figures from the holotype.



Fig. 2. Basidiomata of Collybia effusa.
A. Surface view of the mature pileus. B. Underside view of the mature pileus.
C. Mature basidiomata on a dead twig of Cryptomeria japonica. D. Close up of the stipe
surface. Scales: A, B = 10 mm; C = 20 mm; D = 3 mm. All figures from the holotype.


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